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Journal of The Lepidopterists' Society
Volume 34 1980 Number 1
Journal of the Lepidopterists' Society 34(1), 1980, 1-19
CHECKLIST OF MONTANA BUTTERFLIES (RHOPALOCERA)
Steve Kohler
Montana Department of Natural Resources and Conservation, Forestry Division, 2705 Spurgin Road, Missoula, Montana 59801
ABSTRACT. A list of 177 species of butterflies (Rhopalocera) known to occur in Montana and the county where each specimen has been collected is compiled from records gleaned from university collections, resident collectors, published literature, non-resident collectors, some major natural history museums, and the author's collection.
This checklist has two basic purposes. The first is to present the latest information available on the species of butterflies occurring in Montana and to give their distributions. The second is to serve as a basis for soliciting additional records from non-resident collectors for inclusion in a larger work on Montana's butterflies, which is currently in preparation. This work will include photographs of all species, detailed distribution maps, collection records, life histories and food-plant information. Anyone who can add additional county records for any species is urged to contact me.
Seventy-three years have passed since the first and only attempt at a comprehensive treatment on Montana butterflies was published by M. J. Elrod in 1906, "The Butterflies of Montana," a 174-page illustrated bulletin. An abridged version of this is Elrod and Masters (1970). Early but restricted treatments are those of W. H. Edwards (1872, 1878, 1882b, 1883), Scudder (1875), and Wiley (1894). Papers treating only a species or group of species are Coolidge (1906, 1909), Clench (1944), Daly (1964), W. H. Edwards (1882a, 1886, 1890, 1894), Field (1936a, 1936b, 1938), McDunnough (1928, 1929), Shepard (1964), Skinner (1893, 1921), Stallings and Turner (1947), and Wright (1922). A manuscript checklist of Montana butterflies was prepared in 1941 by Thomas Rogers (Spokane, Washington) but was never published.
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Journal of the Lepidopterists' Society
Elrod's "Butterflies of Montana," though an excellent treatment for its time, is now out-of-date. The nomenclature is antiquated, and the coverage mainly limited to a few counties where collections had been made at that time (Missoula, Flathead, Gallatin, Park, Custer, Lewis & Clark counties). There are also some errors in identification. Copies of the publication are almost impossible to obtain.
Thomas Rogers' manuscript checklist is also restricted in its coverage, listing mainly records from the northwestern part of the state and repeating many of the Elrod records. The nomenclature used is also out-of-date, and, since it was never published, the list has been available to only a very few people.
Montana is a large state; approximately 540 miles in length, with an average width of 275 miles. It is endowed with great variety of habitat and terrain as well as climate. The highest point is Granite Peak (12,799 ft.) in the Beartooth Range, Park Co. The lowest, 1,820 ft, is where Highway 2 crosses the Montana-Idaho border northwest of Troy in Lincoln Co. The western two-fifths of Montana are mountainous; the main Rocky Mountain chain runs north-south across the state. This western mountainous region has received the most attention in the past. The eastern three-fifths of the state are high, rolling prairie country, interrupted by several small mountain ranges. This region has been collected sporadically, and some counties do not have a single species recorded.
Data for county records were obtained from:
(1) The Elrod collection, University of Montana, Missoula. This also includes a collection made by G. E. Barnes in southern Park Co. and the C. A. Wiley collection from Miles City, Custer Co.
(2) The Montana State University collection, Bozeman. Early work on this collection was done by R. A. Cooley, R. E. Hutchins, E. Koch, and A. D. Hastings. The large collection of C. C. Albright, Great Falls, also came to Montana State University at his death, and another small collection by J. J. McDonald of Helena was donated at his death. Albright, a Great Falls physician, collected extensively in the Little Belt Mountains around Monarch, Cascade Co., where he had a summer home. McDonald collected in the vicinity of Helena and the Big Belt Mountains.
(3) The collection of Glacier National Park at park headquarters, West Glacier, which contains eight drawers of butterflies, mainly collected by J. S. Garth in the 1930's.
(4) Private collections of J. E. Crystal, Plains; F. E. Holley, Hamilton; J. Goosey, Jr., Big Timber; and my extensive collection.
(5) A review of the published literature.
(6) Montana collection records provided by non-resident collectors.
Fig. 1. Map of Montana showing county
4 Journal of the Lepidopterists' Society
Table 1. Number of Montana butterfly species recorded by county.
|
Carbon |
128 |
Dawson |
29 |
|
Gallatin |
115 |
Yellowstone |
28 |
|
Flathead |
113 |
Big Horn |
27 |
|
Missoula |
113 |
Blaine |
27 |
|
Sweet Grass |
102 |
Deer Lodge |
23 |
|
Cascade |
96 |
Musselshell |
22 |
|
Lake |
96 |
Prairie |
19 |
|
Lewis & Clark |
95 |
Rosebud |
17 |
|
Madison |
95 |
Fallon |
15 |
|
Ravalli |
87 |
Hill |
13 |
|
Sanders |
83 |
Teton |
8 |
|
Glacier |
78 |
Toole |
8 |
|
Beaverhead |
77 |
Powder River |
7 |
|
Granite |
71 |
Richland |
7 |
|
Powell |
71 |
Wibaux |
7 |
|
Fergus |
69 |
Valley |
5 |
|
Mineral |
67 |
Liberty |
4 |
|
Custer |
66 |
Pondera |
4 |
|
Stillwater |
66 |
Wheatland |
4 |
|
Chouteau |
63 |
Phillips |
2 |
|
Lincoln |
61 |
Roosevelt |
2 |
|
Jefferson |
60 |
Garfield |
1 |
|
Park |
60 |
Petroleum |
1 |
|
Meagher |
49 |
Sheridan |
1 |
|
Judith Basin |
36 |
Treasure |
1 |
|
Broadwater |
35 |
Carter |
0 |
|
Silver Bow |
35 |
Daniels |
0 |
|
Golden Valley |
33 |
McCone |
0 |
(7) Information obtained from the major natural history museums. Unfortunately, there is considerable material in some of these museums that I have not yet examined. It is hoped that this task will be accomplished before the larger work is published. Museums that have collections containing substantial amounts of Montana material are: Allyn Museum of Entomology, Sarasota; American Museum of Natural History, New York; California Academy of Sciences, San Francisco; Canadian National Collection, Ottawa; Carnegie Museum, Pittsburgh; Los Angeles County Museum of Natural History, Los Angeles; Museum of Comparative Zoology, Harvard University, Cambridge; National Museum of Natural History, Washington, D. C; Pea-body Museum of Natural History, Yale University, New Haven; Texas Memorial Museum, Austin. I would certainly appreciate hearing from curators of collections in other institutions if they have Montana material.
Notes on several species are given in the checklist, especially if the nomenclature of the species is in question. Location of counties is
Volume 34, Number 1
5
shown on the Montana map in Fig. 1. Table 1 shows the number of butterfly species recorded in each county.
County Records of Montana Butterflies Megathymidae
Megathymus Scudder, 1872
1 streckeri leussleri Holland—Custer, Rosebud.
Hesperiidae
Amblyscirtes Scudder, 1872
2 vialis (W. H. Edwards)—Carbon, Flathead, Lake, Lincoln, Mineral, Missoula.
Euphyes Scudder, 1872
3 vestris metacomet (Harris)—Sweet Grass.
Ochlodes Scudder, 1872
4a sylvanoides sylvanoides (Roisduval)—Flathead, Granite, Lake, Lewis & Clark, Lincoln, Mineral, Missoula, Ravalli, Sanders.
4b sylvanoides napa (W. H. Edwards)—Broadwater, Carbon, Cascade, Chouteau, Gallatin, Golden Valley, Musselshell, Stillwater, Sweet Grass, Toole, Yellowstone.
Atrytone Scudder, 1872
5 logan lagus (W. H. Edwards)—Cascade, Custer, Powder River. The American
Museum of Natural History has specimens labeled only "Montana" (Stanford, 1975).
Atalopedes Scudder, 1872
6 campestris (Boisduval)—Specimens are in the American Museum of Natural
History which are labeled only "Montana" (Stanford, 1975).
Polites Scudder, 1872
7 peckius (Kirby)—Carbon, Gallatin, Lincoln, Missoula, Petroleum.
8 sabuleti sabuleti (Boisduval)—The American Museum of Natural History has two
specimens labeled only "Montana" (Stanford, 1975).
9 draco (W. H. Edwards)—Beaverhead, Broadwater, Carbon, Chouteau, Fergus,
Flathead, Gallatin, Glacier, Madison, Powell, Stillwater, Sweet Grass.
10 themistocles (Latreille)—Carbon, Cascade, Custer, Flathead, Gallatin, Lewis &
Clark, Madison, Sweet Grass.
11 origenes rhena (W. H. Edwards)—Gallatin, Madison.
12 mystic ssp.—Carbon, Cascade, Chouteau, Gallatin, Lincoln, Mineral, Missoula,
Stillwater, Sweet Grass. MacNeil (in Howe, 1975) feels that Montana, Idaho, Washington, and British Columbia mystic may represent an unnamed subspecies.
13 sonora utahensis (Skinner)—Carbon, Gallatin. There is a specimen from Lake
View, Montana (no county given) in the U.S. National Museum, as well as additional specimens labeled only "Montana" (Stanford, 1975).
Hesperia Fabricius, 1793
14 uncas uncas W. H. Edwards—Beaverhead, Cascade, Chouteau, Fallon, Flathead,
Gallatin, Golden Valley, Hill, Lake, Madison, Missoula, Toole.
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Journal of the Lepidopterists' Society
15a comma assiniboia (Lyman)—Dawson.
15b comma manitoba (Scudder)—Carbon, Flathead, Lake, Lewis & Clark, Missoula, Park, Powell, Sweet Grass, Toole.
15c comma harpalus (W. H. Edwards)—Beaverhead, Broadwater, Carbon, Cascade, Chouteau, Gallatin, Golden Valley, Madison, Meagher, Missoula, Musselshell, Park, Ravalli, Stillwater, Sweet Grass, Yellowstone.
15d comma oregonia (W. H. Edwards)—Sanders.
15e comma oregonia/harpalus—Flathead, Glacier, Lake, Lincoln, Mineral. Specimens of comma from the northwestern part of the state appear to be blends of oregonia and harpalus.
16 nevada (Scudder)—Beaverhead, Carbon, Fergus, Gallatin, Glacier, Golden Val-
ley, Jefferson, Lake, Lewis & Clark, Missoula, Musselshell, Ravalli, Sweet Grass, Yellowstone.
17 pahaska pahaska Leussler—Carbon, Cascade, Golden Valley, Yellowstone.
18 juba (Scudder)—Gallatin, Lake, Madison, Yellowstone.
19 leonardus pawnee Dodge—Big Horn, Chouteau, Custer, Dawson, Prairie. There
are additional specimens collected by Dodge and Neumogen in the U.S. National Museum labeled only "Montana" (Stanford, 1975).
20 ottoe W. H. Edwards—Custer, Prairie. Additional specimens labeled only "Mon-
tana" are in the U.S. National Museum and the American Museum of Natural History (Stanford, 1975).
Oarisma Scudder, 1872
21 garita garita (Reakirt)—Carbon, Cascade, Chouteau, Custer, Flathead, Gallatin,
Glacier, Golden Valley, Granite, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Powell, Ravalli, Richland, Sanders, Sweet Grass.
Carterocephalus Lederer, 1852
22 palaemon mandan (W. H. Edwards)—Beaverhead, Carbon, Cascade, Flathead,
Gallatin, Glacier, Lake, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Sanders, Sweet Grass.
Pholisora Scudder, 1872
23 catullus (Fabricius)—Carbon, Cascade, Chouteau, Custer, Dawson, Gallatin,
Lewis & Clark, Missoula, Ravalli, Sanders, Sweet Grass, Yellowstone.
Hesperopsis Dyar, 1905
24 libya lena (W. H. Edwards)—Custer, Dawson. Additional specimens labeled only
"Montana" are in the collections of the U.S. National Museum, American Museum of Natural History and the Brooklyn Museum (Stanford, 1975; Field, 1976).
Pyrgus Hiibner, 1816
25 centaureae loki Evans—Carbon, Flathead, Glacier, Lake, Lewis & Clark, Park.
26 ruralis (Boisduval)—Beaverhead, Carbon, Cascade, Fergus, Flathead, Gallatin,
Glacier, Golden Valley, Granite, Lake, Lewis & Clark, Lincoln, Madison, Missoula, Park, Ravalli, Sweet Grass.
27 communis communis (Grote)—Blaine, Broadwater, Carbon, Cascade, Chouteau,
Custer, Dawson, Deer Lodge, Fallon, Flathead, Gallatin, Golden Valley, Jefferson, Lake, Lewis & Clark, Madison, Mineral, Missoula, Musselshell, Park, Prairie, Richland, Sanders.
Erynnis Schrank, 1801
28 icelus (Scudder & Burgess)—Carbon, Fergus, Flathead, Gallatin, Glacier, Jeffer-
son, Lincoln, Madison, Mineral, Missoula, Ravalli, Roosevelt, Sanders, Sweet Grass.
29 persius fredericki H. A. Freeman—Beaverhead, Cascade, Chouteau, Fergus, Gal-
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latin, Granite, Lake, Lewis & Clark, Meagher, Mineral, Missoula, Powell, Ravalli, Sanders, Silver Bow, Sweet Grass.
30 afranius (Lintner)—Beaverhead, Cascade, Fergus, Flathead, Gallatin, Glacier,
Judith Basin, Lake, Lewis & Clark, Madison, Meagher, Missoula, Powell.
31 pacuvius lilius (Dyar)—Flathead, Gallatin, Lake, Sanders, Yellowstone.
Thorybes Scudder, 1872
32 pylades pylades (Scudder)—Cascade, Chouteau, Custer, Flathead, Gallatin, Jef-
ferson, Lincoln, Sweet Grass, Yellowstone.
Epargyreus Hiibner, 1816
33 clarus clarus (Cramer)—Big Horn, Carbon, Cascade, Chouteau, Custer, Flathead,
Gallatin, Lake, Lewis & Clark, Lincoln, Missoula, Sanders, Sweet Grass.
Papilionidae
Parnassius Latreille, 1804
34 clodius gallatinus Stichel—Flathead, Gallatin, Glacier, Madison, Mineral, Mis-
soula, Park, Ravalli. References to altaurus Dyar in Montana represent gallatinus. Ferris (1976b) restricts the range of altaurus to Blaine and Custer counties, Idaho.
35a phoebus smintheus Doubleday—Glacier. Ferris (1976a) treated xanthus Ehrmann and idahoensis Bryk & Eisner, both of which have been referred to Montana, as synonyms of smintheus.
35b phoebus montanulus Bryk & Eisner—Beaverhead, Carbon, Cascade, Chouteau, Fergus, Flathead, Gallatin, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Stillwater, Sweet Grass, Teton. Ferris (1976a) has treated maximus Bryk & Eisner as a synonym of montanulus. His arrangement for both smintheus and montanulus seems logical and is followed here, although maximus (TL Judith Mountains, Fergus Co.), because of the large size and melanic appearance of the females, may be distinct enough to be considered a separate subspecies.
Papilio Linnaeus, 1758
36 polyxenes asterius Stoll—Sweet Grass. One specimen, a female, is in the collec-
tion of John Goosey, Jr., Big Timber.
37 bairdii f. "brucei" W. H. Edwards—Carbon, Cascade, Custer, Gallatin, Lewis &
Clark, Madison, Musselshell, Prairie. The taxonomic status of bairdii W. H. Edwards, "brucei," and oregonius W. H. Edwards is still not completely resolved. Emmel's arrangement in Howe (1975) is followed.
38 oregonius W. H. Edwards—Lake, Missoula, Sanders.
39a zelicaon nitra W. H. Edwards—Cascade, Fergus, Glacier, Golden Valley, Granite, Meagher, Missoula, Powell, Sanders. Fisher's (1977) paper outlining his breeding experiments has clarified the relationship of nitra, zelicaon Lucas and "gothica" Remington. His proposed arrangement is followed here.
39b zelicaon nitra f. norm, "gothica"—Beaverhead, Carbon, Cascade, Chouteau, Custer, Deer Lodge, Fallon, Fergus, Flathead, Gallatin, Golden Valley, Granite, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Sweet Grass. This is the much more abundant, yellow "normal form" of nitra.
40 indra indra Reakirt—Carbon, Gallatin, Lewis & Clark.
41 glaucus canadensis Rothschild & Jordan—Carbon, Cascade, Custer, Flathead,
Glacier, Judith Basin, Lincoln, Sanders. Elrod (1906) reports several specimens in Wiley's Miles City collection. Since Elrod outlines the differences between glaucus and rutulus Lucas in his discussion, I feel that these specimens were in
Journal of the Lepidopterists' Society
fact glaucus and not rutulus. The specimens could not be located in the remnants of the Elrod collection.
42 rutulus rutulus Lucas—Beaverhead, Big Horn, Carbon, Deer Lodge, Flathead,
Gallatin, Glacier, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Park, Powell, Sanders, Sweet Grass.
43 multicaudatus Kirby—Beaverhead, Cascade, Flathead, Gallatin, Jefferson, Lake,
Lewis & Clark, Madison, Mineral, Missoula, Pondera, Powell, Ravalli, Rosebud, Sanders, Sweet Grass.
44 eurymedon Lucas—Big Horn, Carbon, Cascade, Custer, Flathead, Gallatin, Gran-
ite, Jefferson, Lake, Lewis & Clark, Madison, Mineral, Missoula, Powell, Ravalli, Sanders, Sweet Grass.
Pieridae
Neophasia Behr, 1869
45a menapia menapia (Felder & Felder)—Carbon, Fergus.
45b menapia nr. tau (Scudder)—Broadwater, Flathead, Granite, Jefferson, Lake,
Lewis & Clark, Lincoln, Mineral, Missoula, Powell, Ravalli, Sanders, Silver
Bow.
Pieris Schrank, 1801
46 beckerii W. H. Edwards—Beaverhead, Broadwater, Carbon, Flathead, Gallatin,
Granite, Jefferson, Lake, Lewis & Clark, Madison, Mineral, Missoula, Park, Sanders, Sweet Grass, Wheatland.
47 sisymbrii elivata (Barnes & Benjamin)—Beaverhead, Flathead, Gallatin, Granite,
Jefferson, Lake, Madison, Missoula, Powell, Ravalli, Sanders.
48 protodice Boisduval & LeConte—Beaverhead, Carbon, Cascade, Chouteau, Cus-
ter, Deer Lodge, Flathead, Gallatin, Golden Valley, Granite, Lake, Lewis & Clark, Madison, Missoula, Park, Yellowstone. Shapiro's (1976) arrangement of the protodice-occidentalis group is followed.
49 occidentalis occidentalis Reakirt—Beaverhead, Blaine, Broadwater, Carbon,
Cascade, Chouteau, Custer, Dawson, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Ravalli, Sanders, Stillwater, Sweet Grass, Teton, Yellowstone.
50 napi macdunnoughii Remington—Beaverhead, Big Horn, Carbon, Cascade,
Chouteau, Custer, Fergus, Flathead, Gallatin, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Stillwater, Sweet Grass.
51 rapae (Linnaeus)—Beaverhead, Big Horn, Blaine, Broadwater, Carbon, Cascade,
Chouteau, Custer, Flathead, Gallatin, Granite, Lake, Lewis & Clark, Madison, Meagher, Mineral, Missoula, Park, Ravalli, Sanders, Stillwater, Sweet Grass, Yellowstone.
Colias Fabricius, 1807
52a meadii meadii W. H. Edwards—Carbon, Gallatin, Madison, Park, Stillwater,
Sweet Grass. 52b meadii elis Strecker—Flathead, Glacier.
53 eurytheme Boisduval—Cascade, Custer, Flathead, Gallatin, Glacier, Granite,
Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Musselshell, Ravalli, Sanders, Sweet Grass.
54 philodice philodice Godart—Beaverhead, Big Horn, Blaine, Broadwater, Carbon,
Cascade, Chouteau, Custer, Dawson, Deer Lodge, Fallon, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Musselshell, Park, Pondera, Powder River, Powell, Ravalli, Rosebud, Sanders, Silver Bow, Still-
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9
water, Sweet Grass, Teton, Valley, Wibaux, Yellowstone. Included here is er-iphyle W. H. Edwards.
55 interior interior Scudder—Flathead, Gallatin, Granite, Jefferson, Lake, Lewis &
Clark, Lincoln, Madison, Mineral, Missoula, Ravalli, Sanders.
56 gigantea harroweri Klots—Gallatin.
57a alexandra columbiensis Ferris—Flathead, Lake, Lincoln, Mineral, Missoula, Ravalli, Sanders. Many specimens show characters associated with both columbiensis and astraea W. H. Edwards (Ferris, 1973).
57b alexandra astraea W. H. Edwards—Beaverhead, Carbon, Cascade, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Jefferson, Lake, Lewis & Clark, Madison, Meagher, Park, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass, Toole. Male specimens grouped here under astraea show a great deal of variation and are mixed orange to yellow populations. Females are white.
58 pelidne skinneri Barnes—Beaverhead, Carbon, Cascade, Gallatin, Glacier, Gran-
ite, Jefferson, Lewis & Clark, Meagher, Missoula, Powell, Ravalli, Stillwater, Sweet Grass.
59 nastes streckeri Grum-Grshmailo—Flathead, Glacier.
Phoebis Hiibner, 1816
60 sennae eubule (Linnaeus)—Carbon.
Nathalis Boisduval, 1836
61 iole Boisduval—Missoula, Stillwater. There is no breeding population of iole in
Montana. The captures surely are migrants.
Anthocharis Boisduval, Rambur & Graslin, 1833
62 sara julia W. H. Edwards—Beaverhead, Broadwater, Carbon, Cascade, Deer
Lodge, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass.
Euchloe Hiibner, 1816
63 ausonides coloradensis (H. Edwards)—Beaverhead, Big Horn, Carbon, Cascade,
Chouteau, Deer Lodge, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Silver Bow, Sweet Grass. Some authors consider coloradensis a synonym of ausonides Lucas (Howe, 1975). California and western mountain populations differ from Rocky Mountain populations in the appearance of the females. The name coloradensis is used here for Montana specimens, and ausonides is restricted to the more western populations.
64 olympia f. "rosa" (W. H. Edwards)—Cascade, Custer, Fallon, Fergus, Gallatin,
Golden Valley, Hill, Musselshell, Park, Prairie, Sweet Grass, "rosa" is best considered a form rather than a subspecies.
65 hyantis hyantis (W. H. Edwards)—Broadwater, Granite, Jefferson, Madison, Ra-
valli, Sanders.
Riodinidae
Apodemia Felder & Felder, 1865
66 mormo mormo (Felder & Felder)—Prairie, Valley.
Lycaenidae
Harkenclenus dos Passos, 1970 67a titus titus (Fabricius)—Dawson.
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Journal of the Lepidopterists' Society
67b titus immaculosus (Comstock)—Carbon, Cascade, Chouteau, Fergus, Flathead, Glacier, Lewis & Clark, Madison, Mineral, Missoula, Ravalli, Stillwater, Sweet Grass.
Satyrium Scudder, 1876
68 fuliginosum semiluna Klots—Big Horn, Carbon, Gallatin, Meagher, Sweet Grass,
Wheatland.
69 saepium okanagana (McDunnough)—Carbon, Cascade, Flathead, Jefferson,
Lake, Mineral, Missoula, Ravalli, Sanders, Stillwater.
70 liparops aliparops (Michener & dos Passos)—Dawson, Prairie, Stillwater, Sweet
Grass.
71 sylvinus ssp.—Carbon, Cascade, Flathead, Jefferson, Lake, Madison, Mineral,
Missoula, Ravalli, Sanders. S. californica (W. H. Edwards) has been reported from Montana on several occasions. I have not been able to examine all the specimens, but those I have seen appear to be sylvinus. For the present, all records for californica have been included here. There are no native Quercus (oak) in Montana, which is the reported larval food of californica, while Salix (willow), the food of sylvinus is widely distributed.
72 acadica montanensis (Watson & Comstock)—Carbon, Custer, Sweet Grass.
Callophrys Billberg, 1820
73 polios obscurus Ferris & Fisher—Carbon, Cascade, Custer, Fergus, Flathead,
Golden Valley, Granite, Lake, Lewis & Clark, Mineral, Missoula, Ravalli, Sanders, Silver Bow.
74 mossii schryveri (Cross)—Granite, Lake, Mineral, Missoula, Ravalli, Sanders,
Silver Bow.
75 augustinus iroides (Boisduval)—Carbon, Fergus, Flathead, Gallatin, Granite,
Lake, Lewis & Clark, Mineral, Missoula, Park^Powell, Ravalli, Sanders.
76 eryphon eryphon (Boisduval)—Beaverhead, Carbon, Cascade, Custer, Fergus,
Flathead, Gallatin, Glacier, Granite, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Powell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass. Elrod (1906) made reference to two records (Miles City, Custer Co. and Bozeman, Gallatin Co.) for niphon (Hiibner). These records are considered doubtful, and the specimens could not be located. For the present, niphon should not be considered as occurring in Montana.
77 spinetorum (Hewitson)—Beaverhead, Carbon, Cascade, Fergus, Flathead, Gal-
latin, Glacier, Granite, Judith Basin, Lake, Lewis & Clark, Lincoln, Missoula, Powell, Ravalli, Sanders.
78 siva siva (W. H. Edwards)—Big Horn, Broadwater, Cascade, Custer, Dawson,
Deer Lodge, Fallon, Gallatin, Granite, Jefferson, Lake, Lewis & Clark, Madison, Missoula, Park, Powell, Rosebud, Sanders, Silver Bow, Stillwater, Sweet Grass, Yellowstone.
79 byrnei Johnson—Ravalli, Sanders. Johnson (1976) has split the nelsoni (Bois-
duval) complex into four different species. Montana specimens from near Nox-on, Sanders Co., were taken closest geographically to the range he outlines for byrnei, though he studied no Montana specimens. Johnson's study was based on relatively few specimens. It remains to be seen if his is the best approach to the nelsoni complex, but his treatment will be followed for the present. Montana specimens may also represent his rosneri, though too few are available to place them accurately.
80 affinis affinis (W. H. Edwards)—Carbon, Fergus, Gallatin, Lewis & Clark, Mad-
ison, Missoula, Sweet Grass.
81 sheridanii nr. neoperplexa (Barnes & Benjamin)—Beaverhead, Big Horn, Car-
bon, Gallatin, Granite, Jefferson, Madison, Powell, Ravalli, Sanders, Silver Bow. Specimens are intermediate between neoperplexa and sheridanii (W. H. Edwards), but appear to be closer to neoperplexa.
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Strymon Hiibner, 1818
82 melinus setonia McDunnough—Big Horn, Carbon, Custer, Dawson, Deer Lodge, Fallon, Fergus, Flathead, Jefferson, Lake, Lewis & Clark, Mineral, Missoula, Ravalli, Rosebud, Sanders, Silver Bow, Stillwater, Sweet Grass, Yellowstone.
Lycaena Fabricius, 1807
83a phlaeas americana Harris—Custer. This subspecies is included on the basis of specimens from Miles City reported by Wiley (Elrod, 1906). These specimens could not be located.
83b phlaeas arethusa (Wolley-Dod)—Flathead, Glacier.
83c phlaeas arctodon Ferris—Carbon, Judith Basin, Sweet Grass.
84 cupreus snowi (W. H. Edwards)—Beaverhead, Carbon, Cascade, Fergus, Flat-
head, Gallatin, Glacier, Lake, Madison, Park, Sweet Grass. Miller and Brown (1979) retain only phlaeas and cupreus in Lycaena. The other species formerly listed in Lycaena have been placed by them in the genera which follow. Their entire arrangement is used here.
Gaeides Scudder, 1876
85 xanthoides dione (Scudder)—Carbon, Cascade, Chouteau, Custer, Dawson, Gla-
cier, Lewis & Clark, Prairie, Stillwater, Sweet Grass.
86 editha montana (Field)—Beaverhead, Broadwater, Carbon, Cascade, Chouteau,
Flathead, Gallatin, Granite, Jefferson, Lake, Lewis & Clark, Madison, Mineral, Missoula, Ravalli, Stillwater, Sweet Grass.
Hyllolycaena L. Miller & F. M. Brown, 1979
87 hyllus (Cramer)—Garfield, Prairie. This species was previously known as thoe
Guerin-Meneville until Brown and Field (1970) showed the correct name as hyllus.
Chalceria Scudder, 1876
88a rubidus sirius (W. H. Edwards)—Blaine, Cascade, Custer, Flathead, Hill, Lake,
Liberty, Powder River. 88b rubidus duofacies (Johnson & Balogh)—Beaverhead, Carbon, Gallatin, Lewis &
Clark, Madison, Park, Powell, Silver Bow, Sweet Grass. This subspecies was
only recently described (Johnson & Balogh, 1977).
89 heteronea klotsi (Field)—Beaverhead, Broadwater, Carbon, Cascade, Flathead,
Gallatin, Glacier, Lake, Lewis & Clark, Madison, Missoula, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass.
Epidemia Scudder, 1876
90 helloides (Boisduval)—Beaverhead, Carbon, Cascade, Chouteau, Custer, Flat-
head, Gallatin, Jefferson, Judith Basin, Lake, Lewis & Clark, Liberty, Madison, Meagher, Missoula, Park, Prairie, Sanders, Stillwater, Sweet Grass, Toole. Ferris (1977) has clarified the relationship of helloides and dorcas (Kirby).
91 dorcas florus (W. H. Edwards)—Beaverhead, Broadwater, Carbon, Cascade,
Chouteau, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Madison, Meagher, Missoula, Park, Ravalli, Sanders, Stillwater, Sweet Grass.
92 mariposa penroseae (Field)—Carbon, Cascade, Flathead, Gallatin, Glacier,
Granite, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Powell, Ravalli, Sanders, Stillwater.
93 nivalis browni (dos Passos)—Beaverhead, Carbon, Flathead, Glacier, Mineral,
Missoula, Ravalli.
Lycaeides Hiibner, 1816 94a argyrognomon atrapraetextus (Field)—Beaverhead, Cascade, Deer Lodge, Flat-
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Journal of the Lepidopterists' Society
head, Glacier, Granite, Lake, Lincoln, Mineral, Missoula, Powell, Ravalli, Sanders. 94b argyrognomon longinus Nabokov—Carbon, Gallatin, Jefferson, Stillwater.
95 melissa melissa (W. H. Edwards)—Beaverhead, Blaine, Broadwater, Carbon,
Cascade, Chouteau, Custer, Dawson, Deer Lodge, Fallon, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Madison, Meagher, Missoula, Musselshell, Park, Powell, Prairie, Ravalli, Richland, Sheridan, Stillwater, Sweet Grass, Teton, Wibaux, Yellowstone.
Plebejus Kluk, 1802
96 saepiolus saepiolus (Boisduval)—Beaverhead, Big Horn, Broadwater, Carbon,
Cascade, Chouteau, Custer, Deer Lodge, Fallon, Fergus, Flathead, Gallatin, Glacier, Judith Basin, Lake, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Richland, Sanders, Silver Bow, Stillwater, Sweet Grass, Wibaux.
97a icarioides lycea (W. H. Edwards)—Beaverhead, Big Horn, Broadwater, Carbon, Cascade, Chouteau, Custer, Deer Lodge, Fallon, Fergus, Gallatin, Jefferson, Madison, Meagher, Park, Silver Bow, Stillwater, Sweet Grass, Yellowstone.
97b icarioides pembina (W. H. Edwards)—Beaverhead, Big Horn, Fallon, Flathead, Glacier, Granite, Lake, Lewis & Clark, Lincoln, Mineral, Missoula, Powell, Ravalli, Sanders.
98 shasta minnehaha (Scudder)—Carbon, Fergus, Meagher, Richland, Sweet Grass.
99 acmon lutzi dos Passos—Carbon, Deer Lodge, Gallatin, Golden Valley, Lewis
& Clark, Lincoln, Madison, Missoula, Park, Ravalli, Sanders, Stillwater, Sweet
Grass. 100a glandon megalo McDunnough—Blaine, Cascade, Chouteau, Fergus, Flathead,
Glacier, Granite, Judith Basin, Lewis & Clark, Lincoln, Missoula, Powell. 100b glandon rustica (W. H. Edwards)—Beaverhead, Carbon, Deer Lodge, Gallatin,
Madison, Park, Stillwater, Sweet Grass.
Everes Hiibner, 1816
101 amyntula albrighti Clench—Blaine, Broadwater, Carbon, Cascade, Chouteau,
Fergus, Flathead, Gallatin, Glacier, Jefferson, Lake, Lincoln, Madison, Missoula, Park, Pondera, Powell, Ravalli, Sanders, Sweet Grass, Teton.
Euphilotes Mattoni, 1977
102 battoides glaucon (W. H. Edwards)—Madison, Missoula, Ravalli. Shields (1975)
discusses placing battoides and enoptes (Boisduval) in Pseudophilotes Beuret, an arrangement followed by several authors including Brown (1972), but retains them in Shijimiaeoides Beuret. Mattoni (1977) more recently erected a new genus, Euphilotes.
103 enoptes ancilla (Barnes & McDunnough)—Carbon, Gallatin, Madison, Missoula,
Ravalli, Sweet Grass.
Glaucopsyche Scudder, 1872
104 lygdamus oro Scudder—Beaverhead, Big Horn, Broadwater, Carbon, Custer,
Deer Lodge, Fallon, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Musselshell, Park, Powell, Ravalli, Rosebud, Sanders, Silver Bow, Stillwater, Sweet Grass, Wibaux, Yellowstone. There is blending with Columbia Skinner in the northwestern counties.
105 piasus daunia W. H. Edwards—Beaverhead, Big Horn, Carbon, Cascade, Deer
Lodge, Fergus, Flathead, Gallatin, Glacier, Granite, Lewis & Clark, Lincoln, Madison, Missoula, Powell, Sanders, Silver Bow, Sweet Grass.
Celastrina Tutt, 1906
106 argiolus pseudargiolus (Boisduval & LeConte)—Big Horn, Carbon, Chouteau,
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13
Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Mineral, Missoula, Ravalli, Sanders, Sweet Grass.
Nymphalidae
Limenitis Fabricius, 1807
107 arthemis rubrofasciata (Barnes & McDunnough)—Flathead, Glacier.
108 archippus archippus (Cramer)—Carbon, Chouteau, Custer, Gallatin, Granite,
Lake, Madison, Missoula, Park, Powell, Ravalli, Sweet Grass, Yellowstone.
109a weidemeyerii latifascia Perkins & Perkins—Beaverhead, Carbon, Cascade, Chouteau, Fergus, Flathead, Gallatin, Granite, Lewis & Clark, Madison, Mineral, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass.
109b weidemeyerii oberfoelli Brown—Custer, Dawson, Prairie.
110 lorquini burrisonii Maynard—Beaverhead, Flathead, Glacier, Granite, Lake,
Lewis & Clark, Lincoln, Mineral, Missoula, Powell, Ravalli, Sanders.
Vanessa Fabricius, 1807
111 atalanta rubria (Fruhstorfer)—Beaverhead, Carbon, Cascade, Chouteau, Daw-
son, Fergus, Flathead, Gallatin, Golden Valley, Judith Basin, Lake, Lewis & Clark, Liberty, Lincoln, Madison, Missoula, Powell, Sanders, Sweet Grass.
112 virginiensis (Drury)—Carbon.
113 cardui (Linnaeus)—Beaverhead, Broadwater, Carbon, Cascade, Chouteau, Cus-
ter, Deer Lodge, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Musselshell, Powell, Ravalli, Rosebud, Sanders, Silver Bow, Stillwater, Sweet Grass, Teton.
114 annabella (Field)—Beaverhead, Carbon, Cascade, Custer, Flathead, Gallatin,
Granite, Lake, Madison, Missoula, Ravalli, Sanders.
Nymphalis Kluk, 1802
115 vau-album watsoni (Hall)—Carbon, Custer, Flathead, Gallatin, Glacier, Lake,
Mineral, Missoula, Ravalli, Sanders, Sweet Grass.
116 californica herri Field—Carbon, Custer, Flathead, Gallatin, Granite, Lake, Lew-
is & Clark, Lincoln, Madison, Mineral, Missoula, Powell, Ravalli, Sanders.
117 milberti furcillata (Say)—Beaverhead, Big Horn, Blaine, Carbon, Cascade,
Chouteau, Custer, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Musselshell, Park, Powell, Prairie, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass.
118 antiopa antiopa (Linnaeus)—Carbon, Cascade, Chouteau, Custer, Fergus, Flat-
head, Gallatin, Glacier, Granite, Hill, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Missoula, Musselshell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass.
Polygonia Hiibner, 1816
119 satyrus satyrus (W. H. Edwards)—Beaverhead, Carbon, Cascade, Chouteau, Fer-
gus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Missoula, Park, Powell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass.
120 faunus rusticus (W. H. Edwards)—Carbon, Cascade, Fergus, Flathead, Gallatin,
Glacier, Granite, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass.
121 zephyrus (W. H. Edwards)—Beaverhead, Blaine, Broadwater, Carbon, Cascade,
Chouteau, Custer, Fergus, Flathead, Gallatin, Glacier, Granite, Judith Basin, Lake, Lewis & Clark, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass.
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Journal of the Lepidopterists' Society
122 oreas silenus (W. H. Edwards)—Carbon, Cascade, Flathead, Lake, Missoula,
Park, Sanders, Sweet Grass. The Carbon, Park and Sweet Grass County records may be doubtful. I have not seen the specimens.
123 progne (Cramer)—Carbon. This also may be a doubtful record, but is retained
for the present, until the specimen can be examined.
Charidryas Scudder, 1872
124 nycteis drusius (W. H. Edwards)—Carbon.
125 gorgone carlotta (Reakirt)—Cascade, Custer, Dawson, Fallon, Fergus, Gallatin,
Phillips, Prairie, Richland, Rosebud, Stillwater, Wibaux, Yellowstone.
126 acastus acastus (W. H. Edwards)—Cascade, Custer, Dawson, Fallon, Flathead,
Gallatin, Lewis & Clark, Prairie.
127 damoetas damoetas (Skinner)—Carbon, Flathead, Glacier.
128 palla calydon (Holland)—Broadwater, Carbon, Cascade, Chouteau, Custer, Fer-
gus, Flathead, Gallatin, Glacier, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Powell, Ravalli, Sanders, Stillwater, Sweet Grass. Some Montana material is near sterope (W. H. Edwards).
Phyciodes Hiibner, 1816
129 tharos ssp.—Beaverhead, Blaine, Broadwater, Carbon, Chouteau, Custer, Dawson, Fergus, Flathead, Gallatin, Glacier, Granite, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Musselshell, Powell, Prairie, Ravalli, Rosebud, Sanders, Stillwater, Sweet Grass.
130 campestris campestris (Behr)—Beaverhead, Blaine, Broadwater, Carbon, Cas-
cade, Chouteau, Custer, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Rosebud, Sanders, Stillwater, Sweet Grass, Yellowstone. There may be blending with camillus (W. H. Edwards) in the southwestern part of the state.
131 mylitta mylitta (W. H. Edwards)—Beaverhead, Big Horn, Blaine, Chouteau,
Flathead, Gallatin, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Missoula, Powell, Ravalli, Sanders, Silver Bow, Sweet Grass.
132 pallida barnesi Skinner—Chouteau, Gallatin, Madison, Ravalli, Sweet Grass.
Hypodryas Higgins, 1978
133 gillettii (Barnes)—Beaverhead, Cascade, Fergus, Glacier, Golden Valley, Judith
Basin, Madison, Mineral, Missoula, Sanders. Higgins (1978) has revised the genus Euphydryas Scudder, and erected two new genera for the North American species. He retained only phaeton (Drury), which does not occur in Montana, in the genus Euphydryas. His arrangement is followed here, though some may prefer to consider his new names as subgeneric, rather than generic.
Occidryas Higgins, 1978
134 colon wallacensis (Gunder)—Beaverhead, Flathead, Lake, Lewis & Clark, Lin-
coln, Mineral, Missoula, Powell, Ravalli, Sanders.
135a anicia anicia (Doubleday)—Flathead, Glacier, Lake, Lincoln, Mineral, Sanders.
135b anicia howlandi (Stallings & Turner)—Beaverhead, Gallatin, Jefferson, Lewis & Clark, Madison, Missoula, Ravalli.
135c anicia bernadetta (Leussler)—Big Horn, Carbon, Cascade, Chouteau, Custer, Fergus, Meagher, Rosebud, Stillwater, Sweet Grass.
136a editha hutchinsi (McDunnough)—Beaverhead, Broadwater, Carbon, Cascade, Flathead, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sweet Grass. Form "montanus" (McDunnough) occurs in the high mountains of the Beartooth Plateau in Carbon and Park counties.
136b editha nr. beani (Skinner)—Flathead.
Volume 34, Number 1
15
Boloria Moore, 1900
137a selene tollandensis (Barnes & Benjamin)—Beaverhead.
137b selene atrocostalis/tollandensis—Beaverhead, Carbon, Cascade, Chouteau, Custer, Dawson, Fergus, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Missoula, Park, Powell, Ravalli, Silver Bow, Sweet Grass. Almost all of Montana, with the exception of a small portion of Beaverhead Co., falls within a broad blend zone of atrocostalis (Huard) and tollandensis (Kohler, 1977). Material from Custer and Dawson counties may prove to be sabulocollis Kohler, but I have not been able to examine the specimens.
138 bellona nr. jenistai Stallings & Turner—Fergus, Flathead, Lake, Lewis & Clark,
Missoula. Montana representatives of bellona are in need of further study.
139 kriemhild (Strecker)—Carbon, Gallatin, Sweet Grass.
140 epithore borealis Perkins—Beaverhead, Flathead, Glacier, Granite, Hill, Lake,
Lewis & Clark, Lincoln, Mineral, Missoula, Powell, Ravalli, Sanders, Silver Bow.
141 freija browni Higgins—Carbon.
142 alberta (W. H. Edwards)—Flathead, Glacier.
143 astarte astarte (Doubleday)—Flathead, Glacier.
144 titania ssp.—Carbon, Cascade, Flathead, Gallatin, Judith Basin, Madison,
Meagher, Missoula, Park, Stillwater, Sweet Grass. The name ingens (Barnes & McDunnough) has been applied to Montana titania, but specimens do not compare well with Wyoming material. Possibly two unnamed subspecies of titania are present in Montana. The species is currently under study by Lee D. Miller.
145 eunomia ursadentis Ferris & Groothuis—Carbon, Stillwater.
Speyeria Scudder, 1872
146 idalia (Drury)—Custer. So far, only a single specimen has been taken, a female
at Miles City by Wiley in 1893.
147 edwardsii (Reakirt)—Blaine, Carbon, Cascade, Chouteau, Custer, Fergus, Flat-
head, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Judith Basin, Lewis & Clark, Madison, Meagher, Missoula, Musselshell, Park, Powder River, Powell, Rosebud, Sanders, Stillwater, Sweet Grass, Valley.
148 coronis ssp.—Beaverhead, Carbon, Cascade, Chouteau, Custer, Deer Lodge, Fer-
gus, Golden Valley, Jefferson, Lewis & Clark, Madison, Meagher, Musselshell, Powder River, Powell, Stillwater, Sweet Grass, Yellowstone. The name hal-cyone (W. H. Edwards) has usually been applied to Montana coronis, but Grey (1979) feels they would be better referred to as snyderi (Skinner). Most Montana coronis are greenish in the ventral hind wing discal area as in snyderi, rather than brownish like halcyone.
149 zerene garretti (Gunder)—Beaverhead, Big Horn, Blaine, Broadwater, Carbon,
Cascade, Chouteau, Deer Lodge, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Hill, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Musselshell, Park, Powell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass, Wheatland.
150a callippe gallatini (McDunnough)—Beaverhead, Broadwater, Carbon, Cascade, Gallatin, Jefferson, Lewis & Clark, Madison, Meagher, Missoula, Park, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass.
150b callippe calgariana (McDunnough)—Blaine, Chouteau, Custer, Dawson, Fergus, Flathead, Glacier, Golden Valley, Lake, Liberty, Sanders, Teton, Valley. There is considerable blending of gallatini and calgariana characters over much of the state.
151a egleis macdunnoughi (Gunder)—Beaverhead, Gallatin, Park, Stillwater.
151b egleis albrighti (Gunder)—Carbon, Cascade, Chouteau, Fergus, Golden Valley, Lewis & Clark, Meagher, Sweet Grass.
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Journal of the Lepidopterists' Society
151c egleis ssp.—Flathead, Glacier, Granite, Lake, Missoula, Powell, Ravalli.
152 atlantis ssp.—Beaverhead, Big Horn, Blaine, Carbon, Cascade, Chouteau, Cus-
ter, Fergus, Flathead, Gallatin, Glacier, Granite, Hill, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Stillwater, Sweet Grass, Toole. There is considerable variation in atlantis across the state. The name hutchinsi (Gunder) is conventionally applied to most material, but specimens approaching beani (Barnes & Benjamin) and Helena dos Passos & Grey are often found in the variation (Grey, 1979).
153 hydaspe sakuntala (Skinner)—Beaverhead, Carbon, Cascade, Chouteau, Fergus,
Flathead, Gallatin, Glacier, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Stillwater, Sweet Grass, Toole.
154 mormonia eurynome (W. H. Edwards)—Beaverhead, Big Horn, Blaine, Broad-
water, Carbon, Cascade, Chouteau, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Hill, Jefferson, Judith Basin, Lake, Lewis & Clark, Madison, Meagher, Missoula, Park, Powell, Ravalli, Silver Bow, Stillwater, Sweet Grass, Teton, Wheatland.
155 cybele leto (Behr)—Beaverhead, Big Horn, Blaine, Carbon, Cascade, Chouteau,
Custer, Dawson, Fergus, Flathead, Gallatin, Glacier, Granite, Hill, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Sweet Grass. Custer and Dawson County material may be closer to nominate cybele (Fabricius), but I have not been able to examine specimens.
156a aphrodite manitoba (Chermock & Chermock)—Big Horn, Custer, Dawson, Musselshell, Rosebud.
156b aphrodite Columbia (H. Edwards)—Flathead, Lake, Lincoln, Missoula.
156c aphrodite ethne (Hemming)—Beaverhead, Blaine, Carbon, Cascade, Chouteau, Fergus, Gallatin, Golden Valley, Hill, Jefferson, Madison, Meagher, Park, Stillwater, Sweet Grass, Treasure. There is considerable intergrading of characters of the different subspecies of aphrodite in several areas.
Euptoieta Doubleday, 1848
157 claudia (Cramer)—Blaine, Carbon, Custer, Dawson, Fergus, Gallatin, Golden
Valley, Hill, Madison, Prairie, Richland, Rosebud, Stillwater, Sweet Grass, Yellowstone.
Danaidae
Danaus Kluk, 1802
158 plexippus plexippus (Linnaeus)—Carbon, Cascade, Custer, Gallatin, Jefferson,
Lake, Lewis & Clark, Madison, Missoula, Musselshell, Sanders, Sweet Grass.
Satyridae
Coenonympha Hiibner, 1816
159a ampelos ampelos W. H. Edwards—Cascade, Flathead, Granite, Lake, Lincoln,
Mineral, Missoula, Ravalli, Sanders. 159b ampelos sweadneri Chermock & Chermock—Mineral, Missoula.
160 ochracea ochracea W. H. Edwards—Beaverhead, Big Horn, Blaine, Broadwater,
Carbon, Cascade, Chouteau, Custer, Fergus, Flathead, Gallatin, Granite, Jefferson, Lewis & Clark, Madison, Missoula, Park, Powell, Rosebud, Silver Bow, Stillwater, Sweet Grass, Yellowstone.
161 inornata benjamini McDunnough—Beaverhead, Blaine, Carbon, Cascade,
Chouteau, Custer, Dawson, Fallon, Flathead, Gallatin, Glacier, Granite, Lake,
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17
Lewis & Clark, Madison, Missoula, Pondera, Roosevelt, Rosebud, Stillwater, Toole, Wibaux, Yellowstone.
162 haydenii (W. H. Edwards)—Broadwater, Carbon, Cascade, Gallatin, Jefferson,
Judith Basin, Lewis & Clark, Madison, Meagher, Park, Powell, Sweet Grass.
Neominois Scudder, 1875
163 ridingsii ridingsii (W. H. Edwards)—Beaverhead, Blaine, Broadwater, Carbon,
Chouteau, Dawson, Fergus, Gallatin, Jefferson, Lewis & Clark, Madison, Park, Sweet Grass.
Cercyonis Scudder, 1875
164 pegala ino (Hall)—Beaverhead, Big Horn, Blaine, Carbon, Cascade, Chouteau,
Custer, Dawson, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Hill, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Musselshell, Park, Phillips, Powder River, Powell, Prairie, Ravalli, Sanders, Stillwater, Sweet Grass, Yellowstone.
165 meadii meadii (W. H. Edwards)—Carbon, Custer, Dawson, Musselshell, Yellow-
stone.
166 sthenele paulus (W. H. Edwards)—Beaverhead, Carbon, Mineral. This species
is included on the basis of records from James A. Scott and C. J. Durden. I have not seen the specimens.
167 oetus charon (W. H. Edwards)—Beaverhead, Blaine, Broadwater, Carbon, Cas-
cade, Chouteau, Custer, Dawson, Flathead, Gallatin, Glacier, Granite, Jefferson, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Mineral, Missoula, Park, Powder River, Powell, Prairie, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass, Valley. Included here are records which reported pho-cus (W. H. Edwards).
Oeneis Hiibner, 1816
168 uhleri varuna (W. H. Edwards)—Blaine, Broadwater, Carbon, Cascade, Chou-
teau, Custer, Dawson, Fallon, Fergus, Flathead, Gallatin, Glacier, Golden Valley, Granite, Jefferson, Lewis & Clark, Meagher, Missoula, Musselshell, Powell, Sweet Grass, Wibaux, Yellowstone.
169 chryxus chryxus (Doubleday)—Beaverhead, Carbon, Cascade, Deer Lodge, Fer-
gus, Flathead, Gallatin, Glacier, Granite, Judith Basin, Lake, Lewis & Clark, Lincoln, Madison, Meagher, Missoula, Park, Powell, Ravalli, Sanders, Silver Bow, Sweet Grass.
170 alberta alberta Elwes—Fergus, Glacier, Golden Valley, Hill, Judith Basin.
171 taygete edwardsi dos Passos—Carbon, Stillwater.
172 jutta reducta McDunnough—Beaverhead, Cascade, Jefferson, Judith Basin,
Lewis & Clark, Meagher, Missoula, Powell. The more northern Montana representatives of jutta are placed as reducta for the present, but specimens require further study.
173 melissa beanii Elwes—Carbon, Glacier, Stillwater.
Erebia Dalman, 1816
174 magdalena magdalena Strecker—Carbon.
175 theano ethela W. H. Edwards—Carbon, Park, Stillwater.
176a epipsodea epipsodea Butler—Beaverhead, Broadwater, Carbon, Deer Lodge, Flathead, Gallatin, Glacier, Granite, Jefferson, Lake, Lewis & Clark, Lincoln, Madison, Mineral, Missoula, Park, Powell, Ravalli, Sanders, Silver Bow, Stillwater, Sweet Grass. .
176b epipsodea freemani Ehrlich—Cascade, Chouteau, Fergus, Judith Basin, Meagher.
177 callias callias W. H. Edwards—Carbon.
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Journal of the Lepidopterists' Society
Acknowledgments
Sincerest appreciation is given to the following for their contributions: Norman L. Anderson (Montana State Univ., Bozeman); Jerry N. Bromenshenk; F. Martin Brown; Patrick J. Conway; Jerald E. Dewey (USDA Forest Service, Missoula); J. E. Crystal; Thomas W. Davies (California Academy of Sciences, San Francisco); Julian P. Donahue (Los Angeles Co. Museum of Natural History, Los Angeles); Christopher J. Durden (Texas Memorial Museum, Austin); J. Gordon Edwards (San Jose State Univ., San Jose); J. Donald Eff; Scott L. Ellis; Clifford D. Ferris (Univ. of Wyoming, Laramie); William D. Field (National Museum of Natural History, Washington, D.C.); Michael S. Fisher; Mr. and Mrs. John Goosey; John Goosey, Jr.; L. Paul Grey; Frank E. Holley; Kurt Johnson (City Univ. of New York); Alexander B. Klots; James H. Lowe (Univ. of Montana, Missoula); Clifford J. Martinka (Glacier National Park, West Glacier); Douglas C. Ferguson (National Museum of Natural History, Washington, D.C.); Bill Mc-Guire; Lee D. Miller (Allyn Museum of Entomology, Sarasota); James H. Pepper; Adam Peters; Edwin M. Perkins, Jr. (Univ. of Southern California, Los Angeles); Ted Pike; Dale F. Schweitzer (Peabody Museum, Yale Univ., New Haven); James A. Scott; Jon H. Shepard; Ray E. Stanford.
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---------. 1909. Melitaea gillettei Barnes. Entomol. News 20: 137.
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of adjacent territories, being a fifth annual report of progress. Govt. Printing Office,
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during 1874. Bull. U.S. Geol. & Geog. Sur. Terr. 4: 513-517. ---------. 1882a. Descriptions of two new species of N. American butterflies. Can. Entomol. 14: 2-6. ---------. 1882b. List of butterflies taken by H. K. Morrison in Dacotah and Montana,
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in 1883, by Wm. M. Courtis, M.E. Papilio 3: 157-164. ---------. 1886. Descriptions of new species of butterflies found in the United States.
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---------. 1894. Description of the preparatory stages of Phyciodes carlotta, Reakirt.
Can. Entomol. 26: 3-8. Elrod, M. J. 1906. The Butterflies of Montana, Univ. of Montana Bull. 30: 1-174. --------- & J. H. MASTERS. 1970. The butterflies of Montana, Mid-Cont. Lepid. Ser.
1(17): 1-14. Ferris, C. D. 1973. A revision of the Colias alexandra complex (Pieridae) aided by
ultraviolet reflectance photography with designation of a new subspecies. J. Lepid.
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