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224

Journal of the Lepidopterists' Society

-------------. 1960b. Food plant relations of the diamond-backed moth (Plutella

maculipennis Curt.) II. Sensory regulation of oviposition of the adult female.

Ent. Expl. Appl. 3: 305-314. Shorey, H. H. 1964. Sex pheromones of noctuid moths. II. Mating behavior of

Trichoplusia ni (Lepidoptera: Noctuidae) with special reference to the role of

the sex pheromone. Ann. Ent. Soc. Amer. 57: 371-377. Thorsteinson, A. J. 1960. Host plant selection in phytophagous insects. Ann. Rev.

Ent. 5: 193-218.

A BILATERAL GYNANDROMORPH OF PIERIS RAPAE (PIERIDAE)

A bilateral gynandromorph of the Imported Cabbage Butterfly, Pieris rapae Linnaeus, was taken September 18, 1969 at the Ithaca, New York city dump (Tompkins Co.). It turned up in a large, randomly collected sample of P. rapae taken for other purposes, and was not recognized as a gynandromorph until already dead. There is thus no information available on its behavior. Nearly all of the butterflies in the sample were taken in flight or on blossoms of Great Burdock (Arctium lappa Linnaeus ).

The specimen is female on the left side and male on the right, similar to one taken in Bedfordshire, England in 1938 (S. H. Kershaw, Proc. S. Lond. ent. not. Hist. Soc. 1954-55, p. 33). All secondary sexual color and pattern characteristics, including the pteridine pigments, are normally developed on the half of appropriate sex. There is no irregular mosaicism. The external genitalia are also bilaterally asymmetrical, with a somewhat aborted clasper on the male side. The internal anatomy was not studied.

A quick survey of the British aberrational literature suggests that gynandromorphs are much rarer in P. rapae than in at least some populations of the P. napi complex. Bilateral gynandromorphs also seem to be very rare in P. brassicae Linnaeus, though irregular mosaics are rather frequent. My specimen is the first of its kind to appear among perhaps 75,000 wild and bred Pieris I have examined in recent years, including some 12,000 wild P. rapae from the northeastern United States.

Arthur M. Shapiro, Department of Entomology and Limnology, Cornell University, Ithaca, New York.

A NOTE ON THE SYSTEMATIC POSITION OF PAPILIO ANTIMACHUS

Paul R. Ehrlich

Department of Biological Sciences, Stanford University, Stanford, California

Recently, the Honorable Miriam Rothschild and Professors J. von Euw, and T. Reichstein (1970) have been able to prove the presence of car-denolide heart poisons in Papilio antimachus Drury. P. antimachus is very unusual in its appearance, with very long wings and a pattern which makes it look something like a giant Acraea. The unusual appearance of the butterfly, and the presence of the cardenolides (heart poisons which also