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8 Fourth Annual Meetings Vol.8: nos.1-2
(Abstract) TAXONOMY AND DISTRIBUTION IN THE GENUS PHILOTES by Rudolph H. T. Mattoni
1. The genus Philotes Scudder is characterized in North America by four distinct types of male genitalia, based on the conformation of the valves.
2. The morphology of the European section of this holarctic genus suggests a phylo-genetic relationship of these types.
3. It is suggested that the basic type of valve is of the Glaucopsyche - Phcedrotes type, which is relatively unmodified in the enoptes group of Philotes.
4. P. battoides Behr is a closer relative to the European section of the genus than to P. enoptes Bdv., and is probably of more recent origin.
5. The European section of the genus consists of three species which are more closely related to one another than are any of the North American section, except for the four species of the enoptes complex.
6. P. speciosa H. Edw. is a species of obscure relationships to any members of the subfamily Glaucopsychinas examined, and may be a relict species.
7. P. sonorensis F. & F. is not congeneric with the remainder of the genus as presently conceived, and a new generic name should be erected for all species other than P. sonorensis discussed in this paper.
8. Philotes battoides Behr and P. glaucon Edw. are conspecific, under the name battoides, by priority.
9. Parallel evolution in sympatric populations of P. enoptes and P. battoides in wing pattern convergence may be explained by recombination under selection of common genetic factors.
10. A new arrangement of the "genus" Philotes is proposed, but until a more complete investigation of the entire subfamily is undertaken, it must remain a tentative working hypothesis:
Philotes (TurananaP)
1. P. speciosa H.Edw.: northwestern Mohave Desert, lower San Joaquin Valley,
Calif., U. S. A.
2. P abencerragus group: Mediterranean region
3. P. vicrama group: central Europe to Himalayas
4. P. baton group: west and south Europe
5. P. battoides group
battoides Behr: arctic alpine High Sierra
oregonensis B. & McD.: Pacific northwest
intermedia B. & McD.: northern California, west side of Sierras
glaucon Edw.: east side of Sierra Nevada, Modoc to Tulare Co.
bernardino B. & McD.: southern Sierra, western Mohave to coast, to northern
Baja California centralis B. & McD.: southern Rockies, Utah, Colorado, no. Arizona.
6. P. enoptes group
enoptes Bdv.: Sierra Nevada, western Nevada, no. Calif.
dammersi Comst. & Henne: Colorado and Mohave deserts to central Arizona
ancilla B. & McD.: Rockies, Montana to New Mexico
7. P. mohave Wats. & Comst.: central Mohave and Colorado deserts
8. P. spaldingi B & McD.: western Rockies
western Calif, to southeastern Arizona
Div. of Botany, University of California, Los Angeles 24, Calif., U. S. A.